100 years ago, titans suddenly appeared on Earth. Soon, human civilization veered on collapse due to the titans. Humans then built a giant wall to defend themselves. Within the giant walls, humans lived in peace, but, 100 years later, the giant wall is broken.
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High school delinquent Seiji Sawamura is desperate to have a girlfriend, especially after being rejected by 20 girls as of late. He's afraid that he will end his life with his right hand as his only companion. Apparently, that doesn't change when one day when he wakes up and discovers that his right hand has become a girl named Midori Kasugano, who confesses that she has had a crush on him for the last three years.
In an alternate timeline, Japan was divided after losing World War II: Hokkaido was annexed by "Union" while Honshu and other southern islands were under US sovereignty. A gigantic yet mysterious tower was constructed at Hokkaido and could be seen clearly from Aomori (the northernmost prefecture of Honshu) across Tsugaru Strait. In the summer of 1996, three 9th-graders had made a promise that one day they'll build an aircraft and unravel the tower's mystery, but their project was abandoned after the girl, Sayuri Sawatari, began experiencing sleeping sickness and transferred to Tokyo for better treatment.